Earth’s ancient sea creatures. Two new species, Troglocladodus trimblei and Glikmanius careforum, were found as part of the park’s Paleontological Resources Inventory, giving insight into what existed during a time when the area was an ancient ocean ecosystem.

The find is not only a witness to the dense history hidden in the cave’s limestone walls but also underscores the untapped potential of the fossils of the park. The Mammoth Cave, now famous for its vast system of caverns, previously existed under an ancient seaway linking the continents of eastern North America, Europe, and northern Africa over 325 million years ago.
Troglocladodus trimblei, named after park Superintendent Barclay Trimble, was revealed via an early PRI expedition in 2019 when Trimble himself discovered the first specimen. The species, which means “Cave Branching Tooth,” was named by adult and juvenile teeth and is estimated to have been approximately 10-12 feet long. The fact that such a creature existed in what is now a landlocked park highlights the evolutionary stories that fossils reveal.
Glikmanius careforum, a new species of the genus Glikmanius, is also believed to have been a fierce predator of its time, reaching the same formidable size. The species was named in honor of the Cave Research Foundation, paying tribute to their support in the research conducted at Mammoth Cave. Most notably, the recent finding of a juvenile Glikmanius’s partial set of jaws and gills represents the first of its kind for this ctenacanthus shark genus, offering valuable insight into its anatomy and hunting behavior.
The fact that these top predators have been found in the fossil record means they once preyed upon the near-shore environments that blanketed Kentucky and Alabama, focusing on small sharks, bony fishes, and orthocones-like squid. They had sharp cutting teeth and estimated high speeds, meaning they were perfectly suited to their environment, controlling the marine ecosystem of their era.
Since the Paleontological Resources Inventory started in 2019, more than 70 ancient fish species have been discovered, highlighting Mammoth Cave as an important paleontological location. Led by experts such as J.P. Hodnett, each of these discoveries keeps pouring in, further adding to the complex tale of our Earth’s biological history.
Rick Toomey, a paleontologist and cave guide at Mammoth Cave, was thrilled with the discovery, saying, “Before four or five years ago, we had absolutely no idea that we had all of these exciting sharks, so many that are new to science and that Mammoth Cave would be one of the most important shark localities from about 340 million years ago.”
The buzz surrounding these discoveries is to be believed, and rightly so. With every find, another layer of Earth’s natural history is stripped away, a glimpse into a bygone era. Though the sites where these fossilized sharks have been discovered are limited to research teams, it becomes evident that Mammoth Cave has many more secrets up its sleeve, waiting to be unearthed by the relentless and inquiring gaze of science.

